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<br>
<img height="33" width="178" src="images/issue.gif" alt="In this issue"><br>
<br>
<b>Human Genome<br>
Map Viewer</b></font></p>
<p><b><font size="2" face="Arial,Helvetica,sans-serif"><a href="profile.html">Investigator
Profile:<br>
Eugene V. Koonin</a></font></b></p>
<p><b><font size="2" face="Arial,Helvetica,sans-serif"><a href="mouse.html">Mouse
Genome<br>
Resources</a></font></b></p>
<p><b><font size="2" face="Arial,Helvetica,sans-serif"><a href="unists.html">UniSTS<br>
Integrates Markers</a></font></b></p>
<p><b><font size="2" face="Arial,Helvetica,sans-serif"><a href="genbank.html">GenBank<br>
Mirror Sites</a></font></b></p>
<p><b><font size="2" face="Arial,Helvetica,sans-serif"><a href="blastlab.html">BLAST
Lab</a> </font></b></p>
<p><b><font size="2" face="Arial,Helvetica,sans-serif"><a href="features.html">New
BLAST<br>
Features</a></font> </b></p>
<p><b><font face="Arial, Helvetica, sans-serif" size="2" color="003399"><a href="masthead.html"><span class="subheads"><span class="subheads">Ma<span class="heads">sthead</span></span></span></a>
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<td width="10" valign="left">&nbsp;</td>
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<p><br>
<br>
<font face="Arial, Helvetica, sans-serif" size="3" color="003399"><b>Tour
the Human Genome with Map Viewer</b></font></p>
<p><font face="Times New Roman, Times, serif" size="3" color="#000000"><br>
NCBI is committed to an ongoing program of incorporating new data and
annotation into its human genome resources and producing updated assemblies
on a regular basis. The data used to generate NCBI's assembly of the
human genome include draft and finished sequence deposited in GenBank
by the Human Genome Project sequencing centers, as well as by individual
contributors. New and updated sequence data continues to be submitted
from both sources.<br>
<br>
The Human Genome Map Viewer provides integrated access to the genome
data through a collection of genetic, physical, and sequence maps. Views
of the data range from specific genes to whole genomic regions of interest.<br>
<br>
A total of 7 maps may be chosen for simultaneous display from a set
of 21, including clone and FISH-mapped clone maps, several radiation
hybrid maps, an EST map, and the Genethon and Marshfield genetic maps.
The Map Viewer can display a particular region of the genome centered
on a gene or marker of interest, or a region defined by an arbitrary
base range. <br>
<br>
<a href="#" onClick="CSAction(new Array(/*CMP*/'B74548FE0'));return CSClickReturn();MM_openBrWindow('genome1.html','Figure1','scrollbars=yes,resizable=yes,width=640,height=480')" csclick="B74548FE0"><img src="images/intro_im.gif" width="421" height="379" border="1"></a>
<font size="1" face="Arial, Helvetica, sans-serif"><b><font color="003399" size="2"><br>
Figure 1:</font></b></font><font size="2" color="003399"> Map Viewer
display of six parallel maps for human BRCA1</font><br>
<br>
<br>
Figure 1 shows six selected maps, with the display centered on the 100Kb
area surrounding the BRCA2 gene. The rightmost is called the Master
Map, which shows the greatest detail and provides additional links.
The other maps provide summary information only. <br>
<br>
In this example, the initial search was for the term &#147;brca2&#148;.
Two BRCA2 records were found on chromosome 13; one for the disease associated
with the gene and one for the gene itself. Following the gene link,
the default display showed two maps&#151;a sequence map labeled &#147;Genes_seq&#148;
and a cytogenetic map labeled &#147;Genes_ cyto&#148;. To view the alternative
set of maps shown in Figure 1, the Display Options and Zoom features
were used. Each of the maps is discussed below.<br>
<br>
The Master Map is designated by the user through the Display Options
function and, in this example, it is the sequence map labelled &#147;Genes_
seq&#148;. The exon structure of the BRCA2 gene is shown as a set of
thick lines superimposed upon the thin line that represents the entire
gene.<br>
<br>
First on the left, the Contig map shows the location of BRCA2 on a particular
contig created by the NCBI genome assembly process. The entire contig
may be downloaded from the map page, or viewed directly as a GenBank
contig record. This contig record contains links to all features found
on the contig, such as STSs, SNPs, FISH-mapped clones, annotated genes
and CDSs.<br>
<br>
Second from the left in Figure 1 is the STS map, which indicates the
positions of sequence-tagged sites in the region of BRCA2. Four STS
markers are shown within the range of the BRCA2 gene given on the Genes_seq
map. Next, the Variation map shows many SNPs within the BRCA2 gene,
with a particularly dense clustering in the region of the largest exon
of BRCA2.<br>
<br>
Figure 1 also includes two recent additions to the palette of maps available
in the human Map Viewer. The EST map shows where ESTs align well to
the genomic sequence, with histograms indicating the mapping density.
Including links to UniGene, the EST map can be used to identify undocumented
exons or to identify the prominent splice variants of genes. The GScan
map shows <i>ab initio</i> gene models derived from GenomeScan, a program
related to the popular GenScan gene prediction tool. This map includes
links to the protein similarities, discovered via a blastx search of
the protein databases, that were used to support the gene predictions.<br>
<br>
Returning to the Genes_seq map, the Master Map in this example, two
links to additional information are provided. The BRCA2 link leads to
the corresponding Locus Link record, which provides a complete summary
of information relating to the gene plus links to related resources.
The &#147;sv&#148; link leads to the NCBI Sequence Viewer, which provides
the most detailed sequence-level view of the human genome.<br>
<br>
<a href="#" onClick="CSAction(new Array(/*CMP*/'B74549611'));return CSClickReturn();MM_openBrWindow('genome1.html','Figure1','scrollbars=yes,resizable=yes,width=640,height=480')" csclick="B74549611"><img src="images/intro_image2.gif" width="421" height="664" border="1" alt="Figure 1"></a><br>
<font color="003399" size="2" face="Arial, Helvetica, sans-serif"><b><br>
Figure 2:</b></font><font color="003399" size="2">&nbsp;Sequence-level
view of the human BRCA2 gene.</font><br>
<br>
<br>
The Sequence Viewer display, a small portion of which is shown in Figure
2, shows the location of exons, CDSs, and STS markers along the BRCA2
gene. This example includes the DNA sequence for the initial region
of the gene, showing the first two exons and intervening intron. One
can see that the start codon is located within the second exon and that
translation of this CDS indicates an initial amino acid sequence of
&#147;MPIGS...&#148; for the BRCA2 gene product. Directly following
is the STS marker sts-X95152, which is also visible on the STS map shown
in Figure 1. The ends of the BRCA2 gene are marked by sts-X95152 and
H48122. Note that the BRCA2 gene is encoded on the reverse complement
of contig NT_009984, such that sts-X95152, which is near the beginning
of the gene, is found at the bottom of the Map Viewer display. The Sequence
Viewer also pin-points the location of three SNPs, two in the first
intron and one in the second exon. All three SNPs are found in noncoding
regions.<br>
<br>
Any portion of the sequence shown in a Map Viewer view may be downloaded
for further analysis using the Download/View feature. A tabular report
for the displayed region can also be generated, including the coordinates
of all Map Viewer features shown on the displayed maps, with links to
each feature.<br>
<br>
An advanced search feature is also offered. Search terms can be restricted
to particular maps; specific features such STSs, clones, or contigs;
certain subsets of SNPs (such as SNPs in coding vs. non-coding regions,
or SNPs with various degrees of heterozygosity); or particular chromosomes.
It is also possible to search for several items together so as to generate
a Map View containing several features, or to search a particular region
defined by a set of markers. A syntax such as &#147;H48122 OR sts-X95157
OR brca2&#148; will generate three hits on chromosome 13, which can
be viewed together.<br>
<br>
The latest NCBI human genome assembly can be downloaded from the Genome
side of the FTP site at <a href="ftp://ftp.ncbi.nih.gov/genomes/H_sapiens">ftp.ncbi.nih.gov/genomes/H_sapiens</a>.<br>
<br>
The FTP site includes sequence data for each chromosome, as well as
the mRNA and protein sequences generated by the NCBI annotation project.
The data used by the Map Viewer to display the various integrated maps
is contained within the &#147;maps&#148; sub-directory. A README file
contains more details.&nbsp;<i><font face="Arial, Helvetica, sans-serif" size="2">&#151;VP,
DW</font></i><br>
<br>
<br>
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