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<title>9MU4: Structure of a native Drosophila melanogaster octameric nucleosome</title>
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<table cellspacing="0" id="mmdb_logo_links" cellpadding="0" width="100%"><tr><td><a class="ncbi_home_logo" href="/" alt="NBI HOME Logo">NCBI</a><div class="NLMLogo">National Center for<br/>Biotechnology Information</div></td><td class="titletext" align="left"><!--a href="/Structure/MMDB/mmdb.shtml"--><img border=0 src="/Structure/MMDB/strbanner.jpg" alt="9MU4 - NCBI MMDB Logo"><!--/a--></td><td style="padding-bottom:1em;vertical-align:bottom"><form id="mmdbsearch" method="get" name="mmdbsearch" action="/Structure/pdb/" style="margin:0;padding:0;vertical-align: bottom;"><div class="mmdb-search-holder" style="height:100%;line-height:100%"><input accesskey="S" id="mmdb-search-box" type="search" name="uid" placeholder="Enter PDB ID or MMDB ID" value = "" aria-describedby="mmdb-search-uid" title="Enter PDB ID or MMDB ID"><button id="mmdb-search-button" class="__button mmdb-search-button" style="height:100%;line-height:100%;margin-right:-3px;" type="submit">Search</button></div></form></td></tr></table>
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<section data-section="infor" id="structure-infor-section"><h1 class="structure-title" data-mmdbid="259629" data-pdbid="9MU4" id="structure-title"><a class="link title-pdbid" href="http://www.rcsb.org/pdb/explore/explore.do?structureId=9MU4" style="font-weight:600;margin:0;">9MU4: </a><span>Structure of a native Drosophila melanogaster octameric nucleosome</span></h1>
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<td><a id="pdb_id_link" class="link" href="http://www.rcsb.org/pdb/explore/explore.do?structureId=9MU4" title="PDB ID">9MU4</a><a id="download-pdb-file" class="__button" tabindex="-1" href="https://files.rcsb.org/download/9MU4.pdb" title="Download PDB file" download="9MU4.pdb">Download</a><a class="help-img" href="/Structure/MMDB/docs/mmdb_help.html#MmdbsrvStructureViewDataSetPDBdataset" title="-click to see help"></a></td>
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<tr><td class="H4 property-name">MMDB ID: </td>
<td style="font-weight:bold"><span class="info-item">259629</span> <a class="help-img" href="/Structure/MMDB/docs/mmdb_help.html#MmdbsrvRecordIdentifiers" title="-click to see help"></a></td>
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<tr><td class="H4 property-name">PDB Deposition Date: </td>
<td><span id="depositionterm" style="padding-left: 0.2em;">2025/1/13<a class="help-img" href="/Structure/MMDB/docs/mmdb_help.html#MmdbsrvDeposited" title="-click to see help"></a></span></td>
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<tr><td class="H4 property-name">Updated in MMDB: </td>
<td class="text" nowrap><span id="mmdb_updated" style="padding-left:0.2em">2025/02</span><a class="help-img" href="/Structure/MMDB/docs/mmdb_help.html#MmdbsrvUpdatedInMMDB" title="-click to see help"></a></td>
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<tr><td class="H4 property-name">Experimental Method: </td>
<td class="TEXT" nowrap><div id="expmethod"><span title="electron microscopy" class="info-item">electron microscopy</span> <a class="help-img" href="/Structure/MMDB/docs/mmdb_help.html#MmdbsrvExperimentalMethod" title="-click to see help"></a></div></td>
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<tr><td class="H4 property-name">Resolution: </td>
<td class="TEXT" id="resolution" nowrap>3.29&nbsp; &#197;<a class="help-img" href="/Structure/MMDB/docs/mmdb_help.html#SearchFieldResolution" title="-click to see help"></a></td>
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<tr><td class="H4 property-name">Source Organism: </td>
<td class="text"><div id="taxonomy"><a id="taxword" class="link" href="//www.ncbi.nlm.nih.gov/Taxonomy/Browser/wwwtax.cgi?lvl=0&id=7227" title="Drosophila melanogaster">Drosophila melanogaster</a><a class="help-img" href="/Structure/MMDB/docs/mmdb_help.html#MmdbsrvTaxonomy" title="-click to see help"></a></div></td>
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<tr><td class="H4 property-name">Similar Structures: </td>
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<section class="content_block" data-section="table"><table border="1" class="bucontent"><caption class="butbl_caption">Molecular Components in 9MU4<a class="help-img" href="/Structure/MMDB/docs/mmdb_help.html#MmdbsrvMolecularComponents" title="-click to see help"></a></caption>
<thead><tr><th>Label</th>
<th>Count</th>
<th>Molecule</th>
</tr></thead>
<tbody data-bun="0" data-type="protein" id="unit0_protein_annotation"><tr><td class="butbl_inner_tbl_tit" colspan="3"><div style="float:left">Proteins (8 molecules)</div></td></tr>
<tr data-row="0"><td class="molecule-label-cell"><div class="identical-molecules floatClear" id="chain-list-0-1" style="text-align: center;"><a href="//www.ncbi.nlm.nih.gov/protein/9MU4_c" id="mol-legend-0-1" class="protein-molecule-label-1" data-gi="2915661890" data-molid="1" style="color:#FFFFFF;background-color:#FF00FF" title="-Select icon for molecule information"><div class="protein-table-chain-label">c</div></a><a href="//www.ncbi.nlm.nih.gov/protein/9MU4_g" id="mol-legend-0-2" class="protein-molecule-label-1" data-gi="2915661891" data-molid="2" style="color:#FFFFFF;background-color:#0000FF" title="-Select icon for molecule information"><div class="protein-table-chain-label">g</div></a></div></td>
<td>2</td>
<td class="annotation-cell"><div class="graph-annot"><div class="molecule-name" title="Histone H2A">Histone H2A</div>
<div class="annot-img-block" data-bun="0" data-gmolid="1" data-molids="1,2" id="molgraph-p-0-1"><img alt="Molecule annotation" src="/Structure/mmdb/images/bigLoader.gif"></div>
</div></td>
</tr>
<tr data-row="1"><td class="molecule-label-cell"><div class="identical-molecules floatClear" id="chain-list-0-2" style="text-align: center;"><a href="//www.ncbi.nlm.nih.gov/protein/9MU4_d" id="mol-legend-0-3" class="protein-molecule-label-1" data-gi="2915661892" data-molid="3" style="color:#FFFFFF;background-color:#996633" title="-Select icon for molecule information"><div class="protein-table-chain-label">d</div></a><a href="//www.ncbi.nlm.nih.gov/protein/9MU4_h" id="mol-legend-0-4" class="protein-molecule-label-1" data-gi="2915661893" data-molid="4" style="color:#000000;background-color:#00FF99" title="-Select icon for molecule information"><div class="protein-table-chain-label">h</div></a></div></td>
<td>2</td>
<td class="annotation-cell"><div class="graph-annot"><div class="molecule-name" title="Histone H2B">Histone H2B</div>
<div class="annot-img-block" data-bun="0" data-gmolid="3" data-molids="3,4" id="molgraph-p-0-3"><img alt="Molecule annotation" src="/Structure/mmdb/images/bigLoader.gif"></div>
</div></td>
</tr>
<tr data-row="2"><td class="molecule-label-cell"><div class="identical-molecules floatClear" id="chain-list-0-3" style="text-align: center;"><a href="//www.ncbi.nlm.nih.gov/protein/9MU4_a" id="mol-legend-0-5" class="protein-molecule-label-1" data-gi="2915661894" data-molid="5" style="color:#000000;background-color:#CCCCCC" title="-Select icon for molecule information"><div class="protein-table-chain-label">a</div></a><a href="//www.ncbi.nlm.nih.gov/protein/9MU4_e" id="mol-legend-0-6" class="protein-molecule-label-1" data-gi="2915661895" data-molid="6" style="color:#000000;background-color:#FF9900" title="-Select icon for molecule information"><div class="protein-table-chain-label">e</div></a></div></td>
<td>2</td>
<td class="annotation-cell"><div class="graph-annot"><div class="molecule-name" title="Histone H3">Histone H3</div>
<div class="annot-img-block" data-bun="0" data-gmolid="5" data-molids="5,6" id="molgraph-p-0-5"><img alt="Molecule annotation" src="/Structure/mmdb/images/bigLoader.gif"></div>
</div></td>
</tr>
<tr data-row="3"><td class="molecule-label-cell"><div class="identical-molecules floatClear" id="chain-list-0-4" style="text-align: center;"><a href="//www.ncbi.nlm.nih.gov/protein/9MU4_b" id="mol-legend-0-7" class="protein-molecule-label-1" data-gi="2915661896" data-molid="7" style="color:#000000;background-color:#FF6666" title="-Select icon for molecule information"><div class="protein-table-chain-label">b</div></a><a href="//www.ncbi.nlm.nih.gov/protein/9MU4_f" id="mol-legend-0-8" class="protein-molecule-label-1" data-gi="2915661897" data-molid="8" style="color:#000000;background-color:#00FF00" title="-Select icon for molecule information"><div class="protein-table-chain-label">f</div></a></div></td>
<td>2</td>
<td class="annotation-cell"><div class="graph-annot"><div class="molecule-name" title="Histone H4">Histone H4</div>
<div class="annot-img-block" data-bun="0" data-gmolid="7" data-molids="7,8" id="molgraph-p-0-7"><img alt="Molecule annotation" src="/Structure/mmdb/images/bigLoader.gif"></div>
</div></td>
</tr>
</tbody>
<tbody data-bun="0" data-type="nucleotide" id="unit0_nucleotide_annotation"><tr><td class="butbl_inner_tbl_tit" colspan="3"><div style="float:left">Nucleotides(2 molecules)</div></td></tr>
<tr data-row="0"><td class="molecule-label-cell"><div class="identical-molecules floatClear" id="chain-list-0-5" style="text-align: center;"><a href="//www.ncbi.nlm.nih.gov/nuccore/9MU4_T" id="mol-legend-0-9" class="drna-molecule-label" data-gi="2915661898" data-molid="9" style="color:#000000;background-color:#00FFFF" title="-Select icon for molecule information"><div class="protein-table-chain-label">T</div></a></div></td>
<td>1</td>
<td class="annotation-cell"><div class="graph-annot"><div class="molecule-name" title="DNA (164-mer)">DNA (164-mer)</div>
<div class="annot-img-block" data-bun="0" data-gmolid="9" data-molids="9" id="molgraph-n-0-9"><img alt="Molecule annotation" src="/Structure/mmdb/images/bigLoader.gif"></div>
</div></td>
</tr>
<tr data-row="1"><td class="molecule-label-cell"><div class="identical-molecules floatClear" id="chain-list-0-6" style="text-align: center;"><a href="//www.ncbi.nlm.nih.gov/nuccore/9MU4_N" id="mol-legend-0-10" class="drna-molecule-label" data-gi="2915661899" data-molid="10" style="color:#000000;background-color:#FFFF99" title="-Select icon for molecule information"><div class="protein-table-chain-label">N</div></a></div></td>
<td>1</td>
<td class="annotation-cell"><div class="graph-annot"><div class="molecule-name" title="DNA (164-mer)">DNA (164-mer)</div>
<div class="annot-img-block" data-bun="0" data-gmolid="10" data-molids="10" id="molgraph-n-0-10"><img alt="Molecule annotation" src="/Structure/mmdb/images/bigLoader.gif"></div>
</div></td>
</tr>
</tbody>
</table>
<div style="margin:0.5em 1em;text-align:left; font-size:85%;">&#42; Click molecule labels to explore molecular sequence information.<br/></div>
</section>
</div></div></div>
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<div align="left" class="citing"><hr width="85%"/><div style="font-weight:bold">Citing MMDB</div><div class="citref" style="padding:0.2em;"><img style="border:0;text-align:left;height:15px;width:15px;margin-right:0.2em" src="//pubchem.ncbi.nlm.nih.gov/images/PubMed.gif"> <span style="color:#444"> Madej T, Lanczycki CJ, Zhang D, Thiessen PA, Geer RC, Marchler-Bauer A, Bryant SH. </span> <cite><a class="link" href="//www.ncbi.nlm.nih.gov/pubmed/24319143">"<span style="font-style:italic;"> MMDB and VAST+: tracking structural similarities between macromolecular complexes. <b style="font-size:90%">Nucleic Acids Res. 2014 Jan; 42(Database issue)</b>:D297-303</span></a></cite></div></div>
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