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<div class="search"><div class="search_form"><label for="database" class="offscreen_noflow">Search database</label><select id="database"><optgroup label="Recent"><option value="books">Books</option><option value="pmc">PMC</option><option value="nuccore" data-ac_dict="nuccore">Nucleotide</option><option value="refseq" class="last">RefSeq</option></optgroup><optgroup label="All"><option value="gquery">All Databases</option><option value="assembly" data-ac_dict="assembly">Assembly</option><option value="biocollections">Biocollections</option><option value="bioproject">BioProject</option><option value="biosample">BioSample</option><option value="books">Books</option><option value="clinvar">ClinVar</option><option value="cdd">Conserved Domains</option><option value="gap">dbGaP</option><option value="dbvar">dbVar</option><option value="gene" selected="selected" data-ac_dict="gene">Gene</option><option value="genome" data-ac_dict="genome">Genome</option><option value="gds">GEO DataSets</option><option value="geoprofiles">GEO Profiles</option><option value="gtr">GTR</option><option value="ipg">Identical Protein Groups</option><option value="medgen">MedGen</option><option value="mesh">MeSH</option><option value="nlmcatalog">NLM Catalog</option><option value="nuccore" data-ac_dict="nuccore">Nucleotide</option><option value="omim">OMIM</option><option value="pmc">PMC</option><option value="protein" data-ac_dict="protein">Protein</option><option value="proteinclusters">Protein Clusters</option><option value="protfam">Protein Family Models</option><option value="pcassay">PubChem BioAssay</option><option value="pccompound">PubChem Compound</option><option value="pcsubstance">PubChem Substance</option><option value="pubmed">PubMed</option><option value="snp">SNP</option><option value="sra">SRA</option><option value="structure">Structure</option><option value="taxonomy">Taxonomy</option><option value="toolkit">ToolKit</option><option value="toolkitall">ToolKitAll</option><option value="toolkitbookgh">ToolKitBookgh</option></optgroup></select><div class="nowrap"><label for="term" class="offscreen_noflow" accesskey="/">Search term</label><div class="nowrap"><input type="text" name="term" id="term" title="Search Gene. Use up and down arrows to choose an item from the autocomplete." value="" class="jig-ncbiclearbutton jig-ncbiautocomplete" data-jigconfig="dictionary:'gene',disableUrl:'NcbiSearchBarAutoComplCtrl'" autocomplete="off" data-sbconfig="ds:'no',pjs:'yes',afs:'yes'" /></div><button id="search" type="submit" class="button_search nowrap" cmd="go">Search</button></div></div><ul class="searchlinks inline_list"><set></set><li><a href="/gene/advanced">Advanced</a></li><li class="help"><a id="help" class="jig-ncbihelpwindow" target="ncbihelp" name="help" href="/books/NBK3841/">Help</a></li></ul></div>
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<div class="facet_cont" data-db="gene" data-default_grps="fg_gene-sources,fg_categories,fg_sequence-content,fg_status" data-sd="db:'gene',op:'select',term:'',qk:'2710',linkname:'',idsfromresult:'4980642',extra:''"><ul class="facet" data-filter_id="gene-sources" id="_gene-sources"><li class="filter_grp "><div class="clearfix"><h3>Gene sources</h3><a href="#" data-value_id="" class="clear clear_hide">Clear</a></div><ul><li class="fil_val"><a href="#" data-value_id="genomic">Genomic</a></li></ul></li></ul><ul class="facet" data-filter_id="categories" id="_categories"><li class="filter_grp "><div class="clearfix"><h3>Categories</h3><a href="#" data-value_id="" class="clear clear_hide">Clear</a></div><ul><li class="fil_val"><a href="#" data-value_id="annotated-genes">Annotated genes</a></li><li class="fil_val"><a href="#" data-value_id="gene-type-protein-coding">Protein-coding</a></li></ul></li></ul><ul class="facet" data-filter_id="sequence-content" id="_sequence-content"><li class="filter_grp "><div class="clearfix"><h3>Sequence content</h3><a href="#" data-value_id="" class="clear clear_hide">Clear</a></div><ul><li class="fil_val"><a href="#" data-value_id="refseq">RefSeq</a></li></ul></li></ul><ul class="facet" data-filter_id="status" id="_status"><li class="filter_grp "><div class="clearfix"><h3>Status</h3><a href="#" data-value_id="" class="clear clear_show">Clear</a></div><ul><li class="fil_val selected"><a href="#" data-value_id="current-only">Current</a><span class="fcount">(1)</span></li></ul></li></ul><div id="chr-location-facet-wrapper" data-bid="MCID_67da123b4f3d5e000d89b6dc" data-hid="MCID_67d9f06cb98da1549268b4db" data-qky="2710" class="disabled" style="display:none"><ul class="facet" data-filter_id="chromosomal-region" id="_chromosomal-region" data-of="yes"><li class="filter_grp of_grp"><div class="clearfix"><h3>Chromosome locations</h3><a href="#" data-value_id="" class="clear clear_hide">Clear</a></div><ul><a id="facet_ofchromosomal-region" href="#facet_of_divchromosomal-region" class="jig-ncbipopper" data-jigconfig="triggerPosition: 'middle right', destPosition: 'middle left', hasArrow: true, arrowDirection: 'left', closeEvent : 'click', openEvent: 'click', adjustFit: 'autoAdjust', openAnimation: 'none', closeAnimation: 'none', addCloseButton:true, width: '375px'" data-value_id="fetch_more">more...</a><div id="facet_of_divchromosomal-region" style="display:none;" class="of_popup"><h4>Chromosome locations</h4><label for="organism-filter" style="width:100%;margin-bottom:.2em;" class="">Select an organism by typing or scrolling</label><input type="text" placeholder="Enter text to narrow down the list" id="organism-filter" style="width:98%;" autocomplete="off" /><br /><div id="chr-region-tax-list"><select id="of_chromosomal-region" size="9" class="chr-region-tax-list" style="width:100%;"></select></div><div id="chr-location-info-ctrl" data-disabled="disabled"><div class="chr-field"><label for="chr-name">Chromosome</label><br /><select id="chr-name" class="chr-field" disabled="disabled"></select></div><div class="chr-field"><label for="chr-from">From</label><br /><input type="text" name="chr-from" id="chr-from" disabled="disabled" class="chr-field" /></div><div class="chr-field"><label for="chr-to">To</label><br /><input type="text" name="chr-to" id="chr-to" disabled="disabled" class="chr-field" /></div></div><div class="actions clearfix"><button class="jig-ncbibutton of_add apply-chr" data-jigconfig="'color': 'ui-ncbibutton-blue'">Apply</button></div></div></ul></li></ul></div><ul class="facet facet_reset"><li><a href="#" data-value_id="reset">Clear all</a></li></ul><ul class="facet_tools"><li><a id="more_filter_groups_link" href="#more_filter_groups" class="jig-ncbipopper" data-jigconfig="triggerPosition: 'middle right', destPosition: 'top left', hasArrow: true, arrowDirection: 'left', closeEvent : 'click', openEvent: 'click', adjustFit: 'autoAdjust', openAnimation: 'none', closeAnimation: 'none', addCloseButton:true, width: '180px'">
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<h1 class="title" id="gene-name"><span class="gn" style="padding-right:.5em;">glaA</span> glucan 1,4-alpha-glucosidase glaA-Aspergillus niger [ <em class="tax">Aspergillus niger</em>
]
</h1>
<span class="geneid">Gene ID: 4980642, updated on
22-Feb-2025</span>
</div>
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<div class="rprt-section gene-summary">
<section class="rprt-section-header" id="summary">
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<div class="rprt-section-body">
<div id="summaryDiv" class="section">
<dl id="summaryDl">
<dt class="noline"> Gene symbol </dt>
<dd class="noline">glaA</dd>
<dt> Gene description </dt>
<dd>glucan 1,4-alpha-glucosidase glaA-Aspergillus niger</dd>
<dt>Locus tag</dt>
<dd>An03g06550</dd>
<dt>See related</dt>
<dd>
<a href="https://fungidb.org/fungidb/app/record/gene/An03g06550" data-gblink-text="Ensembl" ref="ordinalpos=1&amp;ncbi_uid=4980642&amp;link_uid=An03g06550">FungiDB:An03g06550;</a>
<a href="https://veupathdb.org/gene/An03g06550" data-gblink-text="Ensembl" ref="ordinalpos=1&amp;ncbi_uid=4980642&amp;link_uid=An03g06550">VEuPathDB:An03g06550</a>
</dd>
<dt>Gene type</dt>
<dd>protein coding</dd>
<dt>RefSeq status</dt>
<dd>PROVISIONAL</dd>
<dt>Organism</dt>
<dd class="tax">
<a href="/Taxonomy/Browser/wwwtax.cgi?id=5061" ref="ordinalpos=1&amp;ncbi_uid=4980642&amp;link_uid=5061">Aspergillus niger</a>
</dd>
<dt>Lineage</dt>
<dd class="tax">Eukaryota; Fungi; Dikarya; Ascomycota; Pezizomycotina; Eurotiomycetes; Eurotiomycetidae; Eurotiales; Aspergillaceae; Aspergillus; Aspergillus subgen. Circumdati</dd>
<dt>Orthologs</dt>
<dd class="ortholog-links">
<a href="https://www.ncbi.nlm.nih.gov/gene/3507095">Aspergillus fumigatus</a>
<a href="/gene/3507095/ortholog/">all</a>
</dd>
<dt style="padding-top:.5em;">
<span class="new-icon">NEW</span>
</dt>
<dd style="padding-top:.5em;">
<strong>Try the new <a href="https://www.ncbi.nlm.nih.gov/datasets/gene/id/4980642/" data-ga-category="summary" data-ga-action="click_datasets_link" data-ga-label="gene_table" title="View selected attributes for an input list of genes and download datasets containing genomic, transcript and protein sequences along with a detailed data report.">Gene table</a></strong>
<br />
<strong>Try the new <a href="https://www.ncbi.nlm.nih.gov/datasets/gene/id/4980642/products/" data-ga-category="summary" data-ga-action="click_datasets_link" data-ga-label="transcript_table" title="View selected attributes for an input list of genes and download datasets containing genomic, transcript and protein sequences along with a detailed data report.">Transcript table</a></strong>
</dd>
</dl>
</div>
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</div>
<div class="rprt-section gene-genomic-context">
<section class="rprt-section-header" id="genomic-context">
<h2 id="header-genomic-context">
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<span class="ui-ncbitoggler-master-text">Genomic context</span>
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</section>
<div class="rprt-section-body">
<div class="gc_cont">
<div class="gt_cont_contents">
<div style="float:right"><span>See glaA in </span><a href="/genome/gdv/?context=gene&amp;acc=4980642" class="link-to-gdv genome-browser-link" data-genome-browser-link-sort="1" data-acc="">Genome Data Viewer</a> </div>
<dl class="dl-chr-info" style="float:left;margin-top:0;">
<dt>Location: </dt>
<dd>
<span style="display:inline-block;">chromosome: VI, map: unlocalized, clone: An03</span>
</dd>
</dl>
</div>
<dl class="dl-chr-info exon-count">
<dt>Exon count: </dt>
<dd>5</dd>
</dl>
</div>
<p class="withnote margin_t2em"><strong>NT_166520.1</strong><img usemap="#nbrMap4980642_145235852" border="0" alt="Genomic Context describing neighboring genes" src="/gutils/geneneighb.fcgi?geneid=4980642&amp;nuc_gi=145235852&amp;n0=An03g06530&amp;n1=An03g06540&amp;n2=glaA&amp;n3=An03g06560&amp;n4=An03g06570" /><map name="nbrMap4980642_145235852" id="nbrMap4980642_145235852">
<area shape="rect" onMouseOver="showtip(this,event,'uncharacterized protein')" onMouseOut="hidetip()" coords="12,23,120,31" alt="Neighboring gene uncharacterized protein" href="/gene/4980640" />
<area shape="rect" onMouseOver="showtip(this,event,'uncharacterized protein')" onMouseOut="hidetip()" coords="97,35,209,43" alt="Neighboring gene uncharacterized protein" href="/gene/4980641" />
<area shape="rect" onMouseOver="showtip(this,event,'uncharacterized protein')" onMouseOut="hidetip()" coords="271,35,375,43" alt="Neighboring gene uncharacterized protein" href="/gene/4980643" />
<area shape="rect" onMouseOver="showtip(this,event,'uncharacterized protein')" onMouseOut="hidetip()" coords="350,23,423,31" alt="Neighboring gene uncharacterized protein" href="/gene/4980644" />
</map>
</p>
</div>
</div>
<div class="rprt-section gene-genomic-regions-transcripts-products">
<section class="rprt-section-header" id="genomic-regions-transcripts-products">
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<div style="text-align:right;">Go to <a href="#reference-sequences">reference sequence details</a></div>
</div>
<div>
<dl class="dl-chr-info" style="margin-top:-1.2em">
<dt>Genomic Sequence: </dt>
<dd id="single-gen-accession" data-accession="NT_166520.1_1480114_1482285">NT_166520.1 </dd>
</dl>
</div>
<div id="NT_166520" class="divAccession">
<div class="seq-viewer">
<p class="note-link">
<strong>Go to nucleotide:</strong>
<a title="Nucleotide Graphic report" href="/nuccore/NT_166520.1?report=graph&amp;from=1479788&amp;to=1482611&amp;strand=true&amp;app_context=Gene" ref="ordinalpos=1&amp;ncbi_uid=4980642&amp;link_uid=NT_166520.1">Graphics</a>
<a title="Nucleotide FASTA report" href="/nuccore/NT_166520.1?report=fasta&amp;from=1480115&amp;to=1482286&amp;strand=true" ref="ordinalpos=1&amp;ncbi_uid=4980642&amp;link_uid=NT_166520.1">FASTA</a>
<a title="Nucleotide GenBank report" href="/nuccore/NT_166520.1?report=genbank&amp;from=1480115&amp;to=1482286&amp;strand=true" ref="ordinalpos=1&amp;ncbi_uid=4980642&amp;link_uid=NT_166520.1">GenBank</a>
</p>
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</div>
<div class="rprt-section gene-bibliography">
<section class="rprt-section-header" id="bibliography"><h2 id="header-bibliography"><a href="#" class="jig-ncbitoggler-open" data-jigconfig="targets: '.gene-bibliography .rprt-section-body'"><span class="ui-ncbitoggler-master-text">Bibliography</span></a></h2><div class="rprt-section-tools"><a href="#top" title="Go to the top of the page" class="gene-top-page"><span>Go to the top of the page</span></a><a href="/books/NBK3841/#EntrezGene.Bibliography" title="Help" class="gene-section-help"><span>Help</span></a></div></section>
<div class="rprt-section-body" style="">
<div class="generif-pubmed">
<h3 id="relatedarticlesinpubmed">
<a href="#" class="jig-ncbitoggler-open">Related articles in PubMed</a>
</h3>
<div class="generefbl-link">
<ol class="generef-link">
<li>
<a href="/pubmed/23237452/" ref="ordinalpos=1&amp;ncbi_uid=4980642&amp;link_uid=23237452">The transcriptomic fingerprint of glucoamylase over-expression in Aspergillus niger.</a>
<span class="pubmed-auth-date"> Kwon MJ, <em>et al</em>. BMC Genomics, 2012 Dec 13. PMID 23237452, <span class="pubmed-pmc-link"><a href="/pmc/articles/PMC3554566/" ref="ordinalpos=0&amp;ncbi_uid=4980642&amp;link_uid=PMC3554566">Free PMC Article</a></span></span>
</li>
<li>
<a href="/pubmed/17259976/" ref="ordinalpos=2&amp;ncbi_uid=4980642&amp;link_uid=17259976">Genome sequencing and analysis of the versatile cell factory Aspergillus niger CBS 513.88.</a>
<span class="pubmed-auth-date"> Pel HJ, <em>et al</em>. Nat Biotechnol, 2007 Feb. PMID 17259976</span>
</li>
<li>
<a href="/pubmed/22301074/" ref="ordinalpos=3&amp;ncbi_uid=4980642&amp;link_uid=22301074">Manual GO annotation of predictive protein signatures: the InterPro approach to GO curation.</a>
<span class="pubmed-auth-date"> Burge S, <em>et al</em>. Database (Oxford), 2012. PMID 22301074, <span class="pubmed-pmc-link"><a href="/pmc/articles/PMC3270475/" ref="ordinalpos=0&amp;ncbi_uid=4980642&amp;link_uid=PMC3270475">Free PMC Article</a></span></span>
</li>
<li>
<a href="/pubmed/30032202/" ref="ordinalpos=4&amp;ncbi_uid=4980642&amp;link_uid=30032202">TreeGrafter: phylogenetic tree-based annotation of proteins with Gene Ontology terms and other annotations.</a>
<span class="pubmed-auth-date"> Tang H, <em>et al</em>. Bioinformatics, 2019 Feb 1. PMID 30032202, <span class="pubmed-pmc-link"><a href="/pmc/articles/PMC6361231/" ref="ordinalpos=0&amp;ncbi_uid=4980642&amp;link_uid=PMC6361231">Free PMC Article</a></span></span>
</li>
</ol>
</div>
</div>
<div class="generif-pubmed generif">
<h3 id="generifsgenereferencesintofunctions">
<a href="#" class="jig-ncbitoggler-open">GeneRIFs: Gene References Into Functions</a>
</h3>
<div>
<a href="/gene/about-generif" class="whatisthis">What's a GeneRIF?</a>
<div class="generefbl-link">
<ol class="generef-link">
<li class="gene-rif-popper">
<a href="/pubmed/23237452" target="_blank" class="gene-rif-popper" ref="ordinalpos=1&amp;ncbi_uid=4980642&amp;link_uid=23237452">transcriptomic fingerprint of glucoamylase over-expression in Aspergillus niger</a>
<div id="gene-popper1" class="gene-rif-popper">Title: The transcriptomic fingerprint of glucoamylase over-expression in Aspergillus niger.</div>
</li>
</ol>
</div>
<div class="submit-genrif">
<span>Submit:</span>
<a href="/gene/submit-generif/?geneId=4980642&amp;sym=glaA&amp;name=&amp;org=Aspergillus niger">New GeneRIF</a>
<a href="/RefSeq/update.cgi?geneid=4980642&amp;symbol=glaA;task=fixpub">Correction</a>
</div>
</div>
</div>
</div>
</div>
<div class="rprt-section gene-general-gene-info">
<section class="rprt-section-header" id="general-gene-info">
<h2 id="header-general-gene-info">
<a href="#" class="jig-ncbitoggler-open" data-jigconfig="targets: '.gene-general-gene-info .rprt-section-body'">
<span class="ui-ncbitoggler-master-text">General gene information</span>
</a>
</h2>
<div class="rprt-section-tools">
<a href="#top" title="Go to the top of the page" class="gene-top-page">
<span>Go to the top of the page</span>
</a>
<a href="/books/NBK3841/#EntrezGene.General_Gene_Information" title="Help" class="gene-section-help">
<span>Help</span>
</a>
</div>
</section>
<div class="rprt-section-body" style="">
<div></div>
<div class="infosec">
<h3 id="homology">
<a class="jig-ncbitoggler-open">Homology</a>
</h3>
<ul class="geneinfo-subsec">
<li>
<a href="/gene/4980642/ortholog/" id="ncbi-ortholog-link">NCBI Orthologs</a>
</li>
</ul>
</div>
<!--<component id="Gene_InfoPathway"/>-->
<div class="gene-ontology infosec">
<h3 id="gene-ontology">
<a class="jig-ncbitoggler-open">
<span>Gene Ontology</span>
</a>
<a target="_blank" href="https://www.ncbi.nlm.nih.gov/genome/annotation_euk/process/#gene_ontology" class="g-ontology-src" title="Provided by RefSeq">Provided by RefSeq</a>
</h3>
<div class="ontology-tables">
<div class="ui-ncbigrid-outer-div">
<div class="ui-ncbigrid-inner-div">
<table class="jig-ncbigrid" data-jigconfig="isPageable: true, pageSize : 25, isSortable: true, columnTypes: ['str', 'str', 'str']">
<thead>
<tr>
<th class="col1">Function</th>
<th style="width:12%;">Evidence Code</th>
<th style="width:10%;">Pubs</th>
</tr>
</thead>
<tbody>
<tr>
<td headers="ont-func">
<a href="http://amigo.geneontology.org/amigo/term/GO:0004339">enables glucan 1,4-alpha-glucosidase activity</a>
</td>
<td headers="ont-evi- ont-evi-code-">
<a class="jig-ncbipopper" href="http://geneontology.org/docs/guide-go-evidence-codes/" data-jigconfig="destSelector: '#evidence-code-1-0', hasArrow: true, openAnimation: 'none', closeAnimation: 'none', adjustFit: 'autoAdjust', triggerPosition: 'center right', destPosition: 'center left'"> IEA</a>
<div id="evidence-code-1-0" class="ecode">Inferred from Electronic Annotation<br /><a href="http://geneontology.org/docs/guide-go-evidence-codes/">more info</a></div>
</td>
<td headers="ont-evi- ont-pub-"><a href="/pubmed/22301074" target="_blank" title="PubMed ID 22301074" ref="ordinalpos=1&amp;ncbi_uid=4980642&amp;link_uid=22301074">PubMed</a> </td>
</tr>
<tr>
<td headers="ont-func">
<a href="http://amigo.geneontology.org/amigo/term/GO:2001070">enables starch binding</a>
</td>
<td headers="ont-evi- ont-evi-code-">
<a class="jig-ncbipopper" href="http://geneontology.org/docs/guide-go-evidence-codes/" data-jigconfig="destSelector: '#evidence-code-2-0', hasArrow: true, openAnimation: 'none', closeAnimation: 'none', adjustFit: 'autoAdjust', triggerPosition: 'center right', destPosition: 'center left'"> IEA</a>
<div id="evidence-code-2-0" class="ecode">Inferred from Electronic Annotation<br /><a href="http://geneontology.org/docs/guide-go-evidence-codes/">more info</a></div>
</td>
<td headers="ont-evi- ont-pub-"><a href="/pubmed/22301074" target="_blank" title="PubMed ID 22301074" ref="ordinalpos=1&amp;ncbi_uid=4980642&amp;link_uid=22301074">PubMed</a> </td>
</tr>
</tbody>
</table>
</div>
</div>
<div class="ui-ncbigrid-outer-div">
<div class="ui-ncbigrid-inner-div">
<table class="jig-ncbigrid" data-jigconfig="isPageable: true, pageSize : 25, isSortable: true, columnTypes: ['str', 'str', 'str']">
<thead>
<tr>
<th class="col1">Process</th>
<th style="width:12%;">Evidence Code</th>
<th style="width:10%;">Pubs</th>
</tr>
</thead>
<tbody>
<tr>
<td headers="ont-comp">
<a href="http://amigo.geneontology.org/amigo/term/GO:0000272">involved_in polysaccharide catabolic process</a>
</td>
<td headers="ont-evi-2 ont-evi-code-2">
<a class="jig-ncbipopper" href="http://geneontology.org/docs/guide-go-evidence-codes/" data-jigconfig="destSelector: '#evidence-code-1-1', hasArrow: true, openAnimation: 'none', closeAnimation: 'none', adjustFit: 'autoAdjust', triggerPosition: 'center right', destPosition: 'center left'"> IEA</a>
<div id="evidence-code-1-1" class="ecode">Inferred from Electronic Annotation<br /><a href="http://geneontology.org/docs/guide-go-evidence-codes/">more info</a></div>
</td>
<td headers="ont-evi-2 ont-pub-2"><a href="/pubmed/22301074" target="_blank" title="PubMed ID 22301074" ref="ordinalpos=1&amp;ncbi_uid=4980642&amp;link_uid=22301074">PubMed</a> </td>
</tr>
</tbody>
</table>
</div>
</div>
<div class="ui-ncbigrid-outer-div">
<div class="ui-ncbigrid-inner-div">
<table class="jig-ncbigrid" data-config="isPageable: true, pageSize : 25, isSortable: true, columnTypes: ['str', 'str', 'str']">
<thead>
<tr>
<th class="col1">Component</th>
<th style="width:12%;">Evidence Code</th>
<th style="width:10%;">Pubs</th>
</tr>
</thead>
<tbody>
<tr>
<td headers="ont-proc">
<a href="http://amigo.geneontology.org/amigo/term/GO:0000324">located_in fungal-type vacuole</a>
</td>
<td headers="ont-evi-3 ont-evi-code-3">
<a class="jig-ncbipopper" href="http://geneontology.org/docs/guide-go-evidence-codes/" data-jigconfig="destSelector: '#evidence-code-1-2', hasArrow: true, openAnimation: 'none', closeAnimation: 'none', adjustFit: 'autoAdjust', triggerPosition: 'center right', destPosition: 'center left'"> IEA</a>
<div id="evidence-code-1-2" class="ecode">Inferred from Electronic Annotation<br /><a href="http://geneontology.org/docs/guide-go-evidence-codes/">more info</a></div>
</td>
<td headers="ont-evi-3 ont-pub-3"><a href="/pubmed/30032202" target="_blank" title="PubMed ID 30032202" ref="ordinalpos=1&amp;ncbi_uid=4980642&amp;link_uid=30032202">PubMed</a> </td>
</tr>
</tbody>
</table>
</div>
</div>
</div>
</div>
</div>
</div>
<div class="rprt-section gene-general-protein-info">
<section class="rprt-section-header" id="general-protein-info">
<h2 id="header-general-protein-info">
<a href="#" class="jig-ncbitoggler-open" data-jigconfig="targets: '.gene-general-protein-info .rprt-section-body'">
<span class="ui-ncbitoggler-master-text">General protein information</span>
</a>
</h2>
<div class="rprt-section-tools">
<a href="#top" title="Go to the top of the page" class="gene-top-page">
<span>Go to the top of the page</span>
</a>
<a href="/books/NBK3841/#EntrezGene.General_Protein_Information" title="Help" class="gene-section-help">
<span>Help</span>
</a>
</div>
</section>
<div class="rprt-section-body">
<dl>
<dt>Preferred Names</dt>
<dd>glucan 1,4-alpha-glucosidase glaA-Aspergillus niger</dd>
</dl>
<dl>
<dt class="heading">XP_001390530.1</dt>
<dd>
<ul class="geneinfo-subsec">
<li>EC <a href="http://enzyme.expasy.org/EC/3.2.1.3">3.2.1.3</a></li>
</ul>
<ul class="geneinfo-subsec">
<li>Gene-ID: glaA;gla;amyG;<br />Mapping: glaA from A. niger is mapped to chromosome VI (LG VI); see list from DSM, PUBMED 8041364.;<br />Remark: a synonym for glaA is amyG.;<br />Remark: glaA is alternative spliced and encodes preproglucoamylase G1 and preproglucoamylase G2.;<br />extracellular/secretion proteins;<br />See PMID 6204865;<br />See PMID 6440004;<br />See PMID 207655;<br />See PMID 9649747;<br />See PMID 1077246 </li>
</ul>
</dd>
</dl>
</div>
</div>
<div class="rprt-section gene-reference-sequences">
<section class="rprt-section-header" id="reference-sequences">
<h2 id="header-reference-sequences">
<a href="#" class="jig-ncbitoggler-open" data-jigconfig="targets: '.gene-reference-sequences .rprt-section-body'">
<span class="ui-ncbitoggler-master-text">NCBI Reference Sequences (RefSeq)</span>
</a>
</h2>
<div class="rprt-section-tools">
<a href="#top" title="Go to the top of the page" class="gene-top-page">
<span>Go to the top of the page</span>
</a>
<a href="/books/NBK3841/#EntrezGene.NCBI_Reference_Sequences_RefS" title="Help" class="gene-section-help">
<span>Help</span>
</a>
</div>
</section>
<div class="rprt-section-body">
<p>
<span class="new-icon">NEW</span>
<strong>Try the new <a href="/datasets/tables/genes/?table_type=transcripts&amp;ids=4980642&amp;utm_source=gene&amp;utm_medium=referral&amp;utm_campaign=refseq" title="View selected attributes for an input list of genes and download datasets containing genomic, transcript and protein sequences along with a detailed data report." data-ga-category="refseq" data-ga-action="click_datasets_link" data-ga-label="transcript_table">Transcript table</a></strong>
</p>
<h3 class="genomerefseqs gene_std_toggler" id="annotatedrefseq-title">
<a class="jig-ncbitoggler-open">Genome Annotation</a>
</h3>
<div class="genomerefseqs_slave">
<p class="desc"> The following sections contain reference sequences that belong to a
specific genome build. <a href="#" id="expl-genomes">Explain</a></p>
<p id="expl-genomes-desc" style="display:none">This section includes genomic Reference
Sequences (RefSeqs) from all assemblies on which this gene is annotated, such as
RefSeqs for chromosomes and scaffolds (contigs) from both reference and alternate
assemblies. Model RNAs and proteins are also reported here.</p>
<h3 id="refseq-sequencedescription-p106347648">Reference assembly</h3>
<h4 id="genomic-p106347648">Genomic </h4>
<ol>
<li>
<p>NT_166520.1 Reference assembly</p>
<dl>
<dt>Range</dt>
<dd class="rs-range">
<span>1480115..1482286 complement</span>
</dd>
<dt>Download</dt>
<dd><a href="/nuccore/NT_166520.1?from=1480115&amp;to=1482286&amp;report=genbank&amp;strand=true" ref="ordinalpos=1&amp;ncbi_uid=4980642&amp;link_uid=NT_166520.1?from=1480115&amp;to=1482286&amp;report=genbank&amp;strand=true">GenBank</a>, <a href="/nuccore/NT_166520.1?from=1480115&amp;to=1482286&amp;report=fasta&amp;strand=true" ref="ordinalpos=1&amp;ncbi_uid=4980642&amp;link_uid=NT_166520.1?from=1480115&amp;to=1482286&amp;report=fasta&amp;strand=true">FASTA</a>, <a href="/nuccore/NT_166520.1?from=1479788&amp;to=1482611&amp;report=graph&amp;content=5&amp;strand=true" ref="ordinalpos=1&amp;ncbi_uid=4980642&amp;link_uid=NT_166520.1?from=1479788&amp;to=1482611&amp;report=graph&amp;content=5&amp;strand=true">Sequence Viewer (Graphics)</a></dd>
</dl>
</li>
</ol>
<h4 id="mrnaandproteins-p106808784">mRNA and Protein(s) </h4>
<ol>
<li>
<p><a href="/nuccore/XM_001390493.3">XM_001390493.3</a> → <a href="/protein/XP_001390530.1">XP_001390530.1</a>  glucan 1,4-alpha-glucosidase glaA-Aspergillus niger [Aspergillus niger]</p>
<p>
<a href="/ipg/XP_001390530.1/" ref="ncbi_uid=4980642">See identical proteins and their annotated locations for XP_001390530.1</a>
</p>
<p style="background:#fff; margin-top:1em;">Status: PROVISIONAL</p>
<dl>
<dt>UniProtKB/Swiss-Prot</dt>
<dd>
<a href="https://www.uniprot.org/uniprot/P69328" ref="itool=Gene_FullReport&amp;PrId=5380&amp;uid=4980642&amp;nlmid=&amp;db=gene">P69328</a>
</dd>
<dt>UniProtKB/TrEMBL</dt>
<dd><a href="https://www.uniprot.org/uniprot/A0A0A1E2J8" ref="itool=Gene_FullReport&amp;PrId=5380&amp;uid=4980642&amp;nlmid=&amp;db=gene">A0A0A1E2J8</a>, <a href="https://www.uniprot.org/uniprot/A0A370CB08" ref="itool=Gene_FullReport&amp;PrId=5380&amp;uid=4980642&amp;nlmid=&amp;db=gene">A0A370CB08</a>, <a href="https://www.uniprot.org/uniprot/A2QHE1" ref="itool=Gene_FullReport&amp;PrId=5380&amp;uid=4980642&amp;nlmid=&amp;db=gene">A2QHE1</a>, <a href="https://www.uniprot.org/uniprot/F1DHX7" ref="itool=Gene_FullReport&amp;PrId=5380&amp;uid=4980642&amp;nlmid=&amp;db=gene">F1DHX7</a>, <a href="https://www.uniprot.org/uniprot/G3Y7U0" ref="itool=Gene_FullReport&amp;PrId=5380&amp;uid=4980642&amp;nlmid=&amp;db=gene">G3Y7U0</a></dd>
<dt class="clear"> Conserved Domains (2) <a href="/Structure/cdd/wrpsb.cgi?seqinput=XP_001390530.1">summary</a></dt>
<dd>
<dl class="level2">
<dt><a href="/Structure/cdd/cddsrv.cgi?uid=99886">cd05811</a><br /><span class="cddlocation">Location:</span><span class="cddfrom">538</span> → <span class="cddto">639</span></dt>
<dd class="clearfix"> CBM20_glucoamylase; Glucoamylase (glucan1,4-alpha-glucosidase), C-terminal CBM20 (carbohydrate-binding module, family 20) domain. Glucoamylases are inverting, exo-acting starch hydrolases that hydrolyze starch and related polysaccharides by releasing the nonreducing end ...</dd>
</dl>
<dl class="level2">
<dt><a href="/Structure/cdd/cddsrv.cgi?uid=366266">pfam00723</a><br /><span class="cddlocation">Location:</span><span class="cddfrom">37</span> → <span class="cddto">451</span></dt>
<dd class="clearfix"> Glyco_hydro_15; Glycosyl hydrolases family 15</dd>
</dl>
</dd>
</dl>
</li>
</ol>
</div>
</div>
</div>
<div class="rprt-section gene-related-sequences">
<section class="rprt-section-header" id="related-sequences">
<h2 id="header-related-sequences">
<a href="#" class="jig-ncbitoggler-open" data-jigconfig="targets: '.gene-related-sequences .rprt-section-body'">
<span class="ui-ncbitoggler-master-text">Related sequences</span>
</a>
</h2>
<div class="rprt-section-tools">
<a href="#top" title="Go to the top of the page" class="gene-top-page">
<span>Go to the top of the page</span>
</a>
<a href="/books/NBK3841/#EntrezGene.Related_Sequences" title="Help" class="gene-section-help">
<span>Help</span>
</a>
</div>
</section>
<div class="rprt-section-body">
<div>
<table id="proteinTblId" class="jig-ncbiservergrid" data-jigconfig="webservice:'/gene/4980642/data/related-sequences/',webserviceParams: 'p$l=Ajax', isServersideLoad:true, isServersidePaging: true,isServersidePagingCached: true,pageSize:25">
<thead>
<tr>
<th colspan="2" id="rs-nucleo">Nucleotide</th>
<th rowspan="2" id="rs-protein">Protein</th>
</tr>
<tr>
<th id="rs-heading">Heading</th>
<th id="rs-heading-acc">Accession and Version</th>
</tr>
</thead>
<tbody></tbody>
</table>
<table id="proteinTblId" class="jig-ncbigrid" data-jigconfig="isPageable:true, pageSize:25">
<thead>
<tr>
<th class="proteinTblTh" rowspan="2" id="rs-prot-acc">Protein Accession</th>
<th class="proteinTblTh" colspan="2" id="rs-link">Links</th>
</tr>
<tr>
<th id="rs-genepept">GenPept Link</th>
<th id="rs-uniprot">UniProtKB Link</th>
</tr>
</thead>
<tbody>
<tr>
<td headers="rs-prot-acc">P69328.1</td>
<td headers="rs-link rs-genepept"></td>
</tr>
</tbody>
</table>
</div>
</div>
</div>
<div class="rprt-section gene-additional-links">
<section class="rprt-section-header" id="additional-links"><h2 id="header-additional-links"><a href="#" class="jig-ncbitoggler-open" data-jigconfig="targets: '.gene-additional-links .rprt-section-body'"><span class="ui-ncbitoggler-master-text">Additional links</span></a></h2><div class="rprt-section-tools"><a href="#top" title="Go to the top of the page" class="gene-top-page"><span>Go to the top of the page</span></a><a href="/books/NBK3841/#EntrezGene.Additional_Links" title="Help" class="gene-section-help"><span>Help</span></a></div></section>
<div class="rprt-section-body" style="">
<h3 class="withnote" id="genelinkout">
<a class="jig-ncbitoggler">Gene LinkOut</a>
</h3><div>
<p>The following <a href="https://www.ncbi.nlm.nih.gov/projects/linkout/index.html">LinkOut</a> resources are supplied by external providers. These providers are responsible for maintaining the links. </p>
<div>
<div class="linkout">
<a class="jig-ncbitoggler-open main">
<span>Molecular Biology Databases</span>
</a>
<ul>
<li><span class="free-rsc"></span>Kyoto Encyclopedia of Genes and Genomes</li>
<ul>
<li>
<a target="_blank" href="http://www.genome.jp/dbget-bin/www_bget?ang:An03g06550" ref="itool=Gene_FullReport&amp;uid=4980642&amp;nlmid=&amp;db=gene&amp;log$=linkoutlink&amp;PrId=5174">
<span>ang:An03g06550</span>
</a>
</li>
</ul>
</ul>
</div>
<div class="linkout">
<a class="jig-ncbitoggler-open main">
<span>Research Materials</span>
</a>
<ul>
<li><span class="order-rsc"></span>GenScript latest version of gene cDNA ORF Clone</li>
<ul>
<li>
<a target="_blank" href="http://www.genscript.com/gene/4980642" ref="itool=Gene_FullReport&amp;uid=4980642&amp;nlmid=&amp;db=gene&amp;log$=linkoutlink&amp;PrId=8012">
<span>GenScript latest version of gene cDNA ORF Clone</span>
</a>
</li>
</ul>
</ul>
</div>
</div>
</div>
<div id="urls-in-genombrowser" style="display:none">
</div>
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</div>
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<h2 class="offscreen_noflow">Supplemental Content</h2>
<div id="gene-sidebar">
<div>
<div class="portlet brieflink dis-table-of-contents">
<div class="portlet_head">
<div class="portlet_title">
<h3>Table of contents</h3>
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<div class="portlet_content">
<ul>
<li class="">
<a href="#summary">Summary</a>
</li>
<li class="">
<a href="#genomic-context">Genomic context</a>
</li>
<li class="">
<a href="#genomic-regions-transcripts-products">Genomic regions, transcripts, and products</a>
</li>
<li class="">
<a href="#bibliography">Bibliography</a>
</li>
<li class="">
<a href="#general-gene-info">General gene information</a>
<div class="subsection-links"><a href="#homology">Homology</a>, <a href="#gene-ontology">Gene Ontology</a></div>
</li>
<li class="">
<a href="#general-protein-info">General protein information</a>
</li>
<li class="">
<a href="#reference-sequences">NCBI Reference Sequences (RefSeq)</a>
</li>
<li class="">
<a href="#related-sequences">Related sequences</a>
</li>
<li class="">
<a href="#additional-links">Additional links</a>
</li>
</ul>
</div>
</div>
<div class="portlet brieflink dis-genomesbrowser brieflink dis-genomesbrowser">
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<h3>Genome Browsers</h3>
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<li>
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</ul>
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<div class="portlet dis-related-info brieflink">
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<div class="portlet_title">
<h3>Related information</h3>
</div>
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<div class="portlet_content DiscoveryDbLinks">
<ul>
<li class="brieflinkpopper">
<a class="brieflinkpopperctrl" href="/pcassay?LinkName=gene_pcassay_alltarget_list&amp;from_uid=4980642" ref="log$=recordlinks">BioAssay by Target (List)</a>
<div class="brieflinkpop offscreen_noflow">BioAssays related to the gene by protein target or RNAi target</div>
</li>
<li class="brieflinkpopper">
<a class="brieflinkpopperctrl" href="/bioproject?LinkName=gene_bioproject&amp;from_uid=4980642" ref="log$=recordlinks">BioProjects</a>
<div class="brieflinkpop offscreen_noflow">BioProjects related to a gene</div>
</li>
<li class="brieflinkpopper">
<a class="brieflinkpopperctrl" href="/biosystems?LinkName=gene_biosystems&amp;from_uid=4980642" ref="log$=recordlinks">BioSystems</a>
<div class="brieflinkpop offscreen_noflow">BioSystems</div>
</li>
<li class="brieflinkpopper">
<a class="brieflinkpopperctrl" href="/cdd?LinkName=gene_cdd&amp;from_uid=4980642" ref="log$=recordlinks">Conserved Domains</a>
<div class="brieflinkpop offscreen_noflow">Related CDD</div>
</li>
<li class="brieflinkpopper">
<a class="brieflinkpopperctrl" href="/pmc?LinkName=gene_pmc&amp;from_uid=4980642" ref="log$=recordlinks">Full text in PMC</a>
<div class="brieflinkpop offscreen_noflow">PMC links</div>
</li>
<li class="brieflinkpopper">
<a class="brieflinkpopperctrl" href="/pmc?LinkName=gene_pmc_nucleotide&amp;from_uid=4980642" ref="log$=recordlinks">Full text in PMC_nucleotide</a>
<div class="brieflinkpop offscreen_noflow">Full text in PubMedCentral identified from shared sequence links</div>
</li>
<li class="brieflinkpopper">
<a class="brieflinkpopperctrl" href="/sparcle?LinkName=gene_sparcle&amp;from_uid=4980642" ref="log$=recordlinks">Functional Class</a>
<div class="brieflinkpop offscreen_noflow">Functional class of the sequence domain architecture</div>
</li>
<li class="brieflinkpopper">
<a class="brieflinkpopperctrl" href="/gene?LinkName=gene_gene_neighbors&amp;from_uid=4980642" ref="log$=recordlinks">Gene neighbors</a>
<div class="brieflinkpop offscreen_noflow">Overlapping genes and two nearest non-overlapping genes on either side</div>
</li>
<li class="brieflinkpopper">
<a class="brieflinkpopperctrl" href="/nuccore?LinkName=gene_nuccore&amp;from_uid=4980642" ref="log$=recordlinks">Nucleotide</a>
<div class="brieflinkpop offscreen_noflow">Link to related Nucleotide entry</div>
</li>
<li class="brieflinkpopper">
<a class="brieflinkpopperctrl" href="/protein?LinkName=gene_protein&amp;from_uid=4980642" ref="log$=recordlinks">Protein</a>
<div class="brieflinkpop offscreen_noflow">Link to related protein entry</div>
</li>
<li class="brieflinkpopper">
<a class="brieflinkpopperctrl" href="/pubmed?LinkName=gene_pubmed&amp;from_uid=4980642" ref="log$=recordlinks">PubMed</a>
<div class="brieflinkpop offscreen_noflow">Link to related PubMed entry</div>
</li>
<li class="brieflinkpopper">
<a class="brieflinkpopperctrl" href="/pubmed?LinkName=gene_pubmed_rif&amp;from_uid=4980642" ref="log$=recordlinks">PubMed (GeneRIF)</a>
<div class="brieflinkpop offscreen_noflow">Link to related PubMed article from GeneRIFs</div>
</li>
<li class="brieflinkpopper">
<a class="brieflinkpopperctrl" href="/pubmed?LinkName=gene_pubmed_pmc_nucleotide&amp;from_uid=4980642" ref="log$=recordlinks">PubMed(nucleotide/PMC)</a>
<div class="brieflinkpop offscreen_noflow">Citations in PubMed identified from shared sequence and PMC links.</div>
</li>
<li class="brieflinkpopper">
<a class="brieflinkpopperctrl" href="/protein?LinkName=gene_protein_refseq&amp;from_uid=4980642" ref="log$=recordlinks">RefSeq Proteins</a>
<div class="brieflinkpop offscreen_noflow">Link to Protein RefSeqs</div>
</li>
<li class="brieflinkpopper">
<a class="brieflinkpopperctrl" href="/nuccore?LinkName=gene_nuccore_refseqrna&amp;from_uid=4980642" ref="log$=recordlinks">RefSeq RNAs</a>
<div class="brieflinkpop offscreen_noflow">Link to Nucleotide RefSeq RNAs</div>
</li>
<li class="brieflinkpopper">
<a class="brieflinkpopperctrl" href="/taxonomy?LinkName=gene_taxonomy&amp;from_uid=4980642" ref="log$=recordlinks">Taxonomy</a>
<div class="brieflinkpop offscreen_noflow">Link to related taxonomy entry</div>
</li>
</ul>
</div>
</div>
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<div class="portlet_head">
<div class="portlet_title">
<h3>Links to other resources</h3>
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<div class="portlet_content">
<ul>
<li>
<a href="https://fungidb.org/fungidb/app/record/gene/An03g06550">FungiDB</a>
</li>
<li>
<a href="https://veupathdb.org/gene/An03g06550">VEuPathDB</a>
</li>
<li>
<a href="http://www.genome.jp/dbget-bin/www_bget?ang:An03g06550">KEGG</a>
</li>
</ul>
</div>
</div>
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<div class="portlet_head">
<div class="portlet_title">
<h3>General information</h3>
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<a href="/pmc/?term=PMC4383897+OR+PMC3013746+OR+PMC1761442+OR+PMC539985">About Gene</a>
</li>
<li>
<a href="/books/NBK3840">FAQ</a>
</li>
<li>
<a href="https://ftp.ncbi.nlm.nih.gov/gene/">FTP site</a>
</li>
<li>
<a href="/books/NBK3841/">Help</a>
</li>
<li>
<a href="/books/NBK3842/">My NCBI help</a>
</li>
<li>
<a href="/books/NBK21085/">NCBI Handbook</a>
</li>
<li>
<a href="/gene/statistics/">Statistics</a>
</li>
</ul>
</div>
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<a href="https://www.ncbi.nlm.nih.gov/BLAST">BLAST</a>
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<a href="https://www.ncbi.nlm.nih.gov/genome">Genome</a>
</li>
<li>
<a href="https://www.ncbi.nlm.nih.gov/bioproject">BioProject</a>
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<li>
<a href="https://www.ncbi.nlm.nih.gov/genome/gdv/">Genome Data Viewer</a>
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<a href="https://www.ncbi.nlm.nih.gov/geo">GEO</a>
</li>
<li>
<a href="https://www.ncbi.nlm.nih.gov/omim">OMIM</a>
</li>
<li>
<a href="https://www.ncbi.nlm.nih.gov/probe">Probe</a>
</li>
<li>
<a href="https://www.ncbi.nlm.nih.gov/RefSeq">RefSeq</a>
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<a href="https://www.ncbi.nlm.nih.gov/RefSeq/update.cgi?geneid=4980642&amp;symbol=glaA">Submit Correction</a>
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<a href="/gene/submit-generif?geneId=4980642&amp;sym=glaA&amp;name=glucan 1,4-alpha-glucosidase glaA-Aspergillus niger&amp;org=Aspergillus niger">Submit GeneRIF</a>
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<a class="htb ralinkpopperctrl" ref="log$=activity&amp;linkpos=1" href="/portal/utils/pageresolver.fcgi?recordid=67da123b84f3725e594ad504">glaA [Aspergillus niger]</a>
<div class="ralinkpop offscreen_noflow">glaA [Aspergillus niger]<div class="brieflinkpopdesc">Gene ID:4980642</div></div>
<div class="tertiary">Gene</div>
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<li class="ra_rcd ralinkpopper two_line">
<a class="htb ralinkpopperctrl" ref="log$=activity&amp;linkpos=2" href="/portal/utils/pageresolver.fcgi?recordid=67da123a67c23b31e0d45e2f">glucan 1,4-alpha-glucosidase glaA-Aspergillus niger [Aspergillus niger]</a>
<div class="ralinkpop offscreen_noflow">glucan 1,4-alpha-glucosidase glaA-Aspergillus niger [Aspergillus niger]<div class="brieflinkpopdesc">gi|145235763|ref|XP_001390530.1|</div></div>
<div class="tertiary">Protein</div>
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<a class="htb ralinkpopperctrl" ref="log$=activity&amp;linkpos=3" href="/portal/utils/pageresolver.fcgi?recordid=67da12342f30673f7b2062ef">CCR4 CCR4-NOT core exoribonuclease subunit CCR4 [Saccharomyces cerevisiae S288C]</a>
<div class="ralinkpop offscreen_noflow">CCR4 CCR4-NOT core exoribonuclease subunit CCR4 [Saccharomyces cerevisiae S288C]<div class="brieflinkpopdesc">Gene ID:851212</div></div>
<div class="tertiary">Gene</div>
</li>
<li class="ra_rcd ralinkpopper two_line">
<a class="htb ralinkpopperctrl" ref="log$=activity&amp;linkpos=4" href="/portal/utils/pageresolver.fcgi?recordid=67da123284f3725e594ab12e">CCR4-NOT core exoribonuclease subunit CCR4 [Saccharomyces cerevisiae S288C]</a>
<div class="ralinkpop offscreen_noflow">CCR4-NOT core exoribonuclease subunit CCR4 [Saccharomyces cerevisiae S288C]<div class="brieflinkpopdesc">gi|6319298|ref|NP_009381.1|</div></div>
<div class="tertiary">Protein</div>
</li>
<li class="ra_rcd ralinkpopper two_line">
<a class="htb ralinkpopperctrl" ref="log$=activity&amp;linkpos=5" href="/portal/utils/pageresolver.fcgi?recordid=67da122d67c23b31e0d42fa2">PAN2 poly(A) specific ribonuclease subunit PAN2 [Homo sapiens]</a>
<div class="ralinkpop offscreen_noflow">PAN2 poly(A) specific ribonuclease subunit PAN2 [Homo sapiens]<div class="brieflinkpopdesc">Gene ID:9924</div></div>
<div class="tertiary">Gene</div>
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